In his prewar years, Arness worked as a courier for a jewelry wholesaler, loading and unloading railway boxcars at the Minneapolis freight-yards, and logging in Pierce, Idaho. Arness wanted to be a naval fighter pilot, but he felt his poor eyesight would bar him. His height of 6 feet 7 inches (2.01 m) ended his hopes, since 6 feet 2 inches (1.88 m) was the limit for aviators. Instead, he was called for the Army and reported to Fort Snelling, Minnesota in March 1943.
Arness served as a rifleman with the U.S. 3rd Infantry Division, and was severely wounded during Operation Shingle, at Anzio, Italy.
According to James Arness – An Autobiography, he landed on Anzio Beachhead on January 21, 1944 as a rifleman with 2nd Platoon, E Company, 2nd Battalion, 7th Regiment of the 3rd Infantry Division. Due to his height, he was the first ordered off his landing craft to determine the depth of the water; it came up to his waist.
On January 29, 1945, having undergone surgery several times, Arness was honorably discharged. His wounds continued to bother him, and in later years Arness suffered from acute leg pain, which even prevented him from mounting a horse.
His decorations include the Bronze Star Medal, Purple Heart; the European-African-Middle Eastern Campaign Medal with three bronze battle stars, the World War II Victory Medal and the Combat Infantryman Badge.
After his discharge, James Arness entered Beloit College in Wisconsin. He began his performing career as a radio announcer in Minnesota in 1945.
Aurness soon began acting, and appeared in films. He began with RKO, which immediately changed his name to "Arness". His film debut was as Loretta Young's (Katie Holstrom) brother, Peter Holstrom, in The Farmer's Daughter (1947).
Though identified with westerns, Arness also appeared in two science fiction films, The Thing from Another World (in which he portrayed the title character) and Them!. He was a close friend of John Wayne and co-starred with him in Big Jim McLain, Hondo, Island in the Sky, and The Sea Chase.
John Wayne was originally offered the starring role in an upcoming TV western drama titled Gunsmoke. Wayne turned down the offer, but strongly recommended Arness for the role.
After Gunsmoke ended, Arness performed in western-themed movies and television series, including How the West Was Won, and in five made-for-television Gunsmoke movies between 1987 and 1994.
An exception was as a big city police officer in a short-lived 1981 series, McClain's Law. His role as Zeb Macahan in How the West Was Won made him into a cult figure in many European countries, as the series has been re-broadcast many times around Europe, where it became more popular than in the states.
To quote Mr Arness, "With Gunsmoke, we had an outstanding quality of writing. The show had been on radio for three years, so they were able to fine-tune the characters. What made us different from other westerns was the fact that Gunsmoke wasn't just action and a lot of shooting; they were character-study shows. They're interesting to watch all these years later".
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